Prof. Jon R Lloyd


AFFILIATION: University of Manchester,
Williamson Centre of Molecular Environmental Science and Research Centre for Radwaste and Decommissioning

A. Williamson Centre of Molecular Environmental Science and RCRD, The University of Manchester, Williamson Building, Oxford Road, Manchester, M13 9PL, UK.

I lead a multidisciplinary geomicrobiology group working at the interface between biology and geology. Our research focuses on the mechanisms of microbial metal reduction, with emphasis on the environmental impact and biotechnological applications of Fe(III)-reducing bacteria. See for more details


Coker, V.S., Telling, N.D., van der Laan, G., Pattrick, RADP, Pearce, C.I., Arenholz, E., Tuna, F., Winpenny, R., Lloyd, J.R. (2009) Harnessing microbial subsurface metal reduction activities to synthesise nanoscale cobalt ferrite with enhanced magnetic properties.  ACS Nano 3, 1922-1928

Belzano, S., Statham, P.J., Pancost, R.D. and Lloyd, J.R. (2009) Role of microbial populations in the release of reduced iron to the water column from marine aggregates. Aquatic Microbial Ecology  54 291-303

Cutting R.S., Coker V.S., Fellowes J.W., Lloyd J.R. and Vaughan D.J. (2009) Mineralogical and morphological constraints on Fe(III) reduction by Geobacter sulfurreducens  Geochimica et Cosmochimica Acta 73 4004-4022

Rowland, H.A.L., Boothman, C., Pancost, R.., Gault, A.G., Polya, D.A. and Lloyd, J.R. (2009) Microbiological transformations of petroleum and metals leading to arsenic release from West Bengal aquifer sediments.  Journal of Environmental Quality 38 1598-1607  

Macaskie, L.E., I.P.Mikheenko, P.Yong, K.Deplanche, A.J.Murray, M.Paterson-Beedle, V.S. Coker, C.I.Pearce, R.A.D. Pattrick, D.Vaughan, G. Van der Laan and J.R.Lloyd (2009) Today’s Wastes, Tomorrow’s Materials for Environmental Protection. Advanced Materials Research 71-73  541-548

Handley, K.M., Hery, M. and Lloyd, J.R (2009) Redox cycling of arsenic by the hydrothermal marine bacterium Marinobacter santoriniensis.  Environmental Microbiology 11 1601-1611

Renshaw, J.C., Law, N., Geissler, A.G., Livens, F.R. and Lloyd, J.R. (2009) Impact of the Fe(III)-reducing bacteria Geobacter sulfurreducens and Shewanella oneidensis on the speciation of plutonium. Biogeochemistry. 94 191-196

Handley, K.M., Hery, M. and Lloyd, J.R. (2009) Marinobacter santoriniensissp. nov., a novel arsenate-respiring and arsenite-oxidizing bacterium isolated from hydrothermal sediment.   International Journal of Systematic and Evolutionary Microbiology 59(Pt 4): 886-92

Handley-Sidhu, S., Worsfold, P.J.,  Boothman, C., Lloyd, J.R., Alvarez, R.,  Livens, F.R., Vaughan, D.J. and Keith-Roach, M. (2009) Corrosion and fate of depleted uranium penetrators under progressively anaerobic conditions in estuarine sediment Environ. Sci and Technology 43 (2), 350-355

Corkhill, C.L., Wincott, P.L., Lloyd, J.R. and Vaughan, D.J. (2008) The oxidative dissolution of arsenopyrite (FeAsS) and enargite (Cu3AsS4) by Leptospirillum ferrooxidans. Geochimica et Cosmochimica Acta 72 5616-5633

Law, N., Ansari, S., Livens, F.R., , Renshaw, J.C. and Lloyd, J.R. (2008) The formation of nano-scale elemental silver particles via enzymatic reduction by Geobacter sulfurreducens Applied and Environmental Microbiology  74 7090–7093 Doi:10.1128/AEM.01069-08

Jarvis, R.M., Law, N., Shadi, I., O’Brien, P., Lloyd, J.R. and Goodacre, R. (2008).  Surface-enhanced Raman scattering from intracellular and extracellular bacterial locations.  Anal. Chem. 80, 6741–6746

Héry, M., A.G. Gault, H.A. L. Rowland, Gavin Lear, D.A. Polya and JR. Lloyd (2008) Molecular and cultivation-dependent analysis of metal-reducing bacteria implicated in arsenic mobilisation in South East Asian aquifers. Applied Geochemistry 23  3215–3223

Coker, V.S., Pearce, C.I., Pattrick, R.A.D.P., van der Laan, G., Telling, N., Charnock, J.M., Arenholz, E. and Lloyd, J.R. (2008) Probing the site occupancies of Co-, Ni-, and Mn-substituted biogenic magnetite using XAS and XMCD. American Mineralogist 93, 1119–1132

Lloyd, J.R., Pearce, C.I., Coker, V.S., Pattrick, R.A.D.P., van der Laan, G., Cutting, R., Vaughan, D.V., Paterson-Beedle, M., Mikheenko, I., P., Yong, P. and Macaskie, L.E. (2008) Biomineralization; linking the fossil record to the production of high value functional materials. Geobiology 6 285-297

Morris, K., F.R. Livens, J.M. Charnock, I.T. Burke, J. M. McBeth, J. D. C. Begg, C. Boothman and J.R. Lloyd (2008) An X-ray absorption study of the fate of technetium in reduced and reoxidised sediments and mineral phases.  Applied Geochemistry. 23 603–617

Hand, V.L., J.R. Lloyd, D.J. Vaughan, M.J. Wilkins and S.J. Boult (2008) Experimental studies of the influence of grain size, oxygen availability and organic carbon availability on bioclogging in porous media.  Environ. Sci and Technology. 42 1485-1491

Coker, V.S., Bell, A.M.T., Pearce, C.I., Pattrick, R.A.D., van der Laan, G. Lloyd, J.R. (2008) Time-resolved synchrotron X-ray powder diffraction study of the formation of magnetite by the Fe(III)-reducing bacterium, Geobacter sulfurreducens. American Mineralogist 93 540–547

Pearce, C.I., V.S. Coker, J.M. Charnock, R.A.D. Pattrick, J. F. W. Mosselmans, N. Law, TJ. Beveridge and J.R. Lloyd (2008) Microbial manufacture of chalcogenide-based nanoparticles via the reduction  of selenite using Veillonella atypica: an in situ EXAFS study.  Nanotechnology 19 155-603

von Canstein, H., Ogawa, J., Shimizu, S. and Lloyd, J.R. (2008) Secretion of flavins by Shewanella species and their role as extracellular redox mediators.  Applied and Environmental Microbiology. 74 615-623